A guilty plea from the owners of a Manhattan antiques store officials say was selling illegal elephant Ivory.

Brothers Irving and Samuel Morano of Metropolitan Fine Arts and Antiques Inc., will now have to forfeit about $4.5 million of ivory seized from their shop.

That includes more than 1,600 carvings.

Officials say it is the largest seizure of illegal elephant ivory in state history.

As part of their sentence, the Moranos will also have to make a $200,000 contribution to two wildlife conservation organizations.